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There are two possible pathways for photodissociation that can be identied for EtPhCbl but only one (path B) is energetically feasible and involves the lengthening of the CoN Im bond through the MLCT region followed by the lengthening of the CoC bond through the LF region

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However, under conditions of stress and inhibition of the electron transport chain, the enzyme works in the opposite direction by transferring electrons from NADPH to NAD + , which impairs mitochondrial antioxidant defenses and feeds more reducing equivalents into the impaired electron transport chain leading to increased electron leakage and superoxide formation [133]