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In addition to 2D cell culture, 3D organoid systems or co-culture systems with stromal fibroblasts, immune cells or endothelial cells have increased translational relevance by modelling the tumour microenvironment, heterotypic signalling and drug penetration kinetics (219, 220)

In contrast, several genes associated with cell plasticity (Tead1, Fzd4 , and Tagln2) and neural crest-like features ( Ngfr and Erbb3 ) were significantly upregulated on FN relative to COL I (Fig